Meier, M. Geschichte der Völkerwanderung. Europa, Asien und Afrika vom 3. bis zum 8. Jahrhundert n. Chr. (C. H. Beck, 2021).
Schmidt-Hofner, S. Barbarian migrations and the economic challenges to the Roman landholding elites in the fourth century CE. J. Late Antiq. 10, 372–404 (2017).
Halsall, G. Barbarian Migrations and the Roman West 376–568 (Cambridge Univ. Press, 2007).
Bavuso, I. & Castrorao Barba, A. in The European Countryside during the Migration Period: Patterns of Change from Iberia to the Caucasus (300–700 CE) (eds Bavuso, I. & Castrorao Barba, A.) 1–11 (De Gruyter, 2023).
Halsall, G. in Fifth-Century Gaul: A Crisis of Identity? (eds Drinkwater, J. & Elton, H.) 196–207 (Cambridge Univ. Press, 1992).
Brownlee, E. Grave goods in Early Medieval Europe: regional variability and decline. Internet Archaeol. https://doi.org/10.11141/ia.56.11 (2021).
Hamerow, H. Early Medieval Settlements: The Archaeology of Rural Communities in Northwest Europe, 400–900 (Oxford Univ. Press, 2004).
Brather-Walter, S. in The European Countryside during the Migration Period: Patterns of Change from Iberia to the Caucasus (300-700 CE) (eds Bavuso, I. & Castrorao Barba, A.) 201–231 (De Gruyter, 2023).
Theuws, F. in The Oxford Handbook of the Merovingian World (eds Effros, B. & Moreira, I.) 883–915 (Oxford Univ. Press, 2020).
Brather, S. in Rome, Constantinople and Newly-Converted Europe. Archaeological and Historical Evidence Vol. 1 (eds Salamon, M. et al.) 333–349 (Kraków, Leipzig, Rzeszów, Warszawa, 2012).
Sebrich, J. Das spätantik-frühmittelalterliche Gräberfeld von Essenbach-Altheim. Materialhefte zur Bayerischen Archäologie 110 (Michael Laßleben, 2019).
Heitmeier, I. & Haberstroh, J. Gründerzeit: Siedlung in Bayern zwischen Spätantike und frühem Mittelalter (EOS, 2019).
Fehr, H. & Heitmeier, I. Die Anfänge Bayerns. Von Raetien und Noricum zur frühmittelalterlichen Baiovaria (EOS, 2012).
Veeramah, K. R. et al. Population genomic analysis of elongated skulls reveals extensive female-biased immigration in Early Medieval Bavaria. Proc. Natl Acad. Sci. USA 115, 3494–3499 (2018).
Amorim, C. E. G. et al. Understanding 6th-century barbarian social organization and migration through paleogenomics. Nat. Commun. 9, 3547 (2018).
Gretzinger, J. et al. The Anglo-Saxon migration and the formation of the early English gene pool. Nature 610, 112–119 (2022).
Patterson, N. et al. Large-scale migration into Britain during the Middle to Late Bronze Age. Nature 601, 588–594 (2022).
Antonio, M. L. et al. Stable population structure in Europe since the Iron Age, despite high mobility. eLife 13, e79714 (2024).
Vyas, D. N. et al. Fine-scale sampling uncovers the complexity of migrations in 5th-6th century Pannonia. Curr. Biol. 33, 3951–3961.e11 (2023).
Olalde, I. et al. A genetic history of the Balkans from Roman frontier to Slavic migrations. Cell 186, 5472–5485.e9 (2023).
O’Sullivan, N. et al. Ancient genome-wide analyses infer kinship structure in an Early Medieval Alemannic graveyard. Sci. Adv. 4, eaao1262 (2018).
Speidel, L. et al. High-resolution genomic history of early medieval Europe. Nature 637, 118–126 (2025).
Speidel, L., Forest, M., Shi, S. & Myers, S. R. A method for genome-wide genealogy estimation for thousands of samples. Nat. Genet. 51, 1321–1329 (2019).
Lawson, D. J., Hellenthal, G., Myers, S. & Falush, D. Inference of population structure using dense haplotype data. PLoS Genet. 8, e1002453 (2012).
de Gennaro, L. et al. PANE: fast and reliable ancestral reconstruction on ancient genotype data with non-negative least square and principal component analysis. Genome Biol. 26, 29 (2025).
Lee, A. D. War in Late Antiquity: A Social History (Blackwell, 2007).
McColl, H. et al. Steppe ancestry in western Eurasia and the spread of the Germanic Languages. Preprint at bioRxiv https://doi.org/10.1101/2024.03.13.584607 (2024).
Velte, M. et al. Between Raetia Secunda and the duchy of Bavaria: exploring patterns of human movement and diet. PLoS ONE 18, e0283243 (2023).
Tian, Y. et al. The role of emerging elites in the formation and development of communities after the fall of the Roman Empire. Proc. Natl Acad. Sci. USA 121, e2317868121 (2024).
Treggiari, S. Roman Marriage: Lusti Coniuges from the Time of Cicero to the Time of Ulpian (Oxford Univ. Press, 1991).
Frier, B. W. in The Cambridge Ancient History: The High Empire, AD 70–192 Vol. 11 (eds Bowman, A. K., Garnsey, P. & Rathbone, D.) 787–816 (Cambridge Univ. Press, 2000).
Barbiera, I., Castiglioni, M. & Dalla-Zuanna, G. Demography, peasantry, and family in Early Medieval Provence, 813–814. Population 77, 249–274 (2022).
Jussen, B. Der Name der Witwe. Erkundungen zur Semantik der mittelalterlichen Bußkultur (Veröffentlichungen des Max-Planck-Instituts für Geschichte 158) (Göttingen, 2000).
Ubl, K. Inzestverbot und Gesetzgebung: Die Konstruktion eines Verbrechens (300–1100) (Millennium-Studien 20) (De Gruyter, 2008).
Evans-Grubbs, J. in A companion to Late Antiquity (ed. Rousseau, P.) 201–219 (Wiley-Blackwell, 2009).
Harper, K. in The Oxford Handbook of Late Antiquity (ed. Johnson, S. F.) 667–714 (Oxford, 2012).
Wang, K. et al. Ancient DNA reveals reproductive barrier despite shared Avar-period culture. Nature 638, 1007–1014 (2025).
Saag, L. et al. North Pontic crossroads: mobility in Ukraine from the Bronze Age to the early modern period. Sci. Adv. 11, eadr0695 (2025).
Gnecchi-Ruscone, G. A. et al. Ancient genomes reveal origin and rapid trans-Eurasian migration of 7th century Avar elites. Cell 185, 1402–1413.e21 (2022).
Maróti, Z. et al. The genetic origin of Huns, Avars, and conquering Hungarians. Curr. Biol. 32, 2858–2870.e7 (2022).
Gretzinger, J. et al. Ancient DNA connects large-scale migration with the spread of Slavs. Nature 646, 384–393 (2025).
Akram-Lodhi, A. H. et al. (eds) Handbook of Critical Agrarian Studies (Edward Elgar, 2021).
Kohl, T. Lokale Gesellschaften: Formen der Gemeinschaft in Bayern vom 8. bis zum 10. Jahrhundert (Mittelalter-Forschungen 29) (Ostfildern, 2010).
Guyon, L., Guez, J., Toupance, B., Heyer, E. & Chaix, R. Patrilineal segmentary systems provide a peaceful explanation for the post-Neolithic Y-chromosome bottleneck. Nat. Commun. 15, 3243 (2024).
Guyon, L., Heyer E., Chaix, R. Was descent in Neolithic and Bronze Age Europe patrilineal or bilateral? Proc. Biol. Sci. 292, 20250815 (2025).
Ly, G. et al. From matrimonial practices to genetic diversity in Southeast Asian populations: the signature of the matrilineal puzzle. Philos. Trans. R. 374, 20180434 (2019).
Arjava, A. Women and Law in Late Antiquity (Oxford Univ. Press, 1996).
Gardner, J. F. Women in Roman Law & Society (Croom Helm, 1986).
Shaw, B. D. & Saller, R. P. Close-kin marriage in Roman society? Man 19, 432–444 (1984).
Cassidy, L. M. et al. Continental influx and pervasive matrilocality in Iron Age Britain. Nature 637, 1136–1142 (2025).
Gnecchi-Ruscone, G. A. et al. Network of large pedigrees reveals social practices of Avar communities. Nature 629, 376–383 (2024).
Rivollat, M. et al. Extensive pedigrees reveal the social organization of a Neolithic community. Nature 620, 600–606 (2023).
Brather, S. Ethnische Interpretationen in der frühgeschichtlichen Archäologie. Geschichte, Grundlagen und Alternativen (Ergänzungsbände zum Reallexikon der germanischen Altertumskunde 42) (Walter de Gruyter, 2004).
Brather, S. in Antike im Mittelalter – Fortleben, Nachwirken, Wahrnehmung. 25 Jahre Forschungsverbund “Archäologie und Geschichte des ersten Jahrtausends in Südwestdeutschland” (eds Brather, S. et al.) 217–234 (Ostfildern, 2014).
Pohl, W. in Strategies of Identification: Ethnicity and Religion in Early Medieval Europe (eds Pohl, W. & Heydemann, G.) 1–64 (Turnhout, 2013).
Schipp, O. Der weströmische Kolonat von Konstantin bis zu den Karolingern (332 bis 861). Studien zur Geschichtsforschung des Altertums (21) (Dr. Kovac, 2009).
Schulz, I. et al. Ancient genomes provide evidence of demographic shift to Slavic-associated groups in Moravia. Genome Biol. 26, 259 (2025).
Haubrichs, W. in The Languages of Early Medieval Charters. Latin, Germanic Vernaculars and the Written Word (eds Gallagher, R. et al.) 68–116 (Brill, 2021).
Zedda, N. et al. Biological and substitute parents in Beaker period adult-child graves. Sci Rep. 13, 18765 (2023).
Link, V. et al. ATLAS: Analysis Tools for Low-depth and Ancient Samples. Preprint at bioRxiv https://doi.org/10.1101/105346 (2017).
Tarasov, A., Vilella, A. J., Cuppen, E., Nijman, I. J. & Prins, P. Sambamba: fast processing of NGS alignment formats. Bioinformatics 31, 2032–2034 (2015).
McKenna, A. et al. The Genome Analysis Toolkit: a MapReduce framework for analyzing next-generation DNA sequencing data. Genome Res. 20, 1297–1303 (2010).
Rubinacci, S., Ribeiro, D. M., Hofmeister, R. J. & Delaneau, O. Efficient phasing and imputation of low-coverage sequencing data using large reference panels. Nat. Genet. 53, 120–126 (2021).
Rubinacci, S., Hofmeister, R., da Mota, B. S. & Delaneau, O. Imputation of low-coverage sequencing data from 150,119 UK Biobank genomes. bioRxiv https://doi.org/10.1101/2022.11.28.518213 (2022).
Mathieson, I. et al. Genome-wide patterns of selection in 230 ancient Eurasians. Nature 528, 499–503 (2015).
Schönherr, S., Weissensteiner, H., Kronenberg, F. & Forer, L. Haplogrep 3—an interactive haplogroup classification and analysis platform. Nucleic Acids Res. 51, W263–W268 (2023).
Ralf, A., González, D. M., Zhong, K. & Kayser, M. Yleaf: software for human Y-chromosomal haplogroup inference from next-generation sequencing data. Mol. Biol. Evol. 35, 1291–1294 (2018).
Mallick, S. et al. The Allen Ancient DNA Resource (AADR) a curated compendium of ancient human genomes. Sci. Data 11, 182 (2024).
Mallick, S. & Reich, D. The Allen Ancient DNA Resource (AADR): a curated compendium of ancient human genomes. Harvard Dataverse https://doi.org/10.7910/DVN/FFIDCW (2023).
Patterson, N., Price, A. L. & Reich, D. Population structure and eigenanalysis. PLoS Genet. 2, e190 (2006).
Maier, R. et al. On the limits of fitting complex models of population history to f-statistics. eLife 12, e85492 (2023).
Maier, R. & Patterson, N. admixtools: Inferring demographic history from genetic data. R package version 2.0.4. GitHub https://github.com/uqrmaie1/admixtools (2024).
1000 Genomes Project Consortium. A global reference for human genetic variation. Nature 526, 68–74 (2015).
Chacón-Duque, J.-C. et al. Latin Americans show wide-spread Converso ancestry and imprint of local Native ancestry on physical appearance. Nat. Commun. 9, 5388 (2018).
Ringbauer, H., Novembre, J. & Steinrücken, M. Parental relatedness through time revealed by runs of homozygosity in ancient DNA. Nat. Commun. 12, 5425 (2021).
Ringbauer, H. et al. Accurate detection of identity-by-descent segments in human ancient DNA. Nat. Genet. 56, 143–151 (2024).
Popli, D., Peyrégne, S. & Peter, B. M. KIN: a method to infer relatedness from low-coverage ancient DNA. Genome Biol. 24, 10 (2023).
Alaçamlı, E. et al. READv2: advanced and user-friendly detection of biological relatedness in archaeogenomics. Genome Biol. 25, 216 (2024).
Blöcher, J. et al. Descent, marriage, and residence practices of a 3,800-year-old pastoral community in Central Eurasia. Proc. Natl Acad. Sci. USA 120, e2303574120 (2023).
Pouyet, F., Aeschbacher, S., Thiéry, A. & Excoffier, L. Background selection and biased gene conversion affect more than 95% of the human genome and bias demographic inferences. eLife 7, e36317 (2018).
Haller, B. C. & Messer, P. W. SLiM 3: forward genetic simulations beyond the Wright-Fisher model. Mol. Biol. Evol. 36, 632–637 (2019).
Seabold, S. & Perktold, J. in Proc. 9th Python in Science Conference (eds van der Walt, S. & Millman, J.) 92–96 (SciPy, 2010).
GEBCO Compilation Group. GEBCO 2022 Grid https://doi.org/10.5285/e0f0bb80-ab44-2739-e053-6c86abc0289c (2022).
Hunter, J. D. Matplotlib: a 2D graphics environment. Comput. Sci. Eng. 9, 90–95 (2007).
Wegmann, D., Eckel, R. Demography and life histories across the Roman frontier in Southern Germany 400–700 ce. Zenodo https://doi.org/10.5281/zenodo.17192653 (2026).

