Botella, H., Blom, H., Dorka, M., Ahlberg, P. E. & Janvier, P. Jaws and teeth of the earliest bony fishes. Nature 448, 583–586 (2007).
Chen, D. L., Blom, H., Sanchez, S., Tafforeau, P. & Ahlberg, P. E. The stem osteichthyan Andreolepis and the origin of tooth replacement. Nature 539, 237–241 (2016).
Chen, D. L. et al. Development of cyclic shedding teeth from semi-shedding teeth: the inner dental arcade of the stem osteichthyan Lophosteus. R. Soc. Open Sci. 4, 161084 (2017).
Chen, D. L. et al. The developmental relationship between teeth and dermal odontodes in the most primitive bony fish Lophosteus. eLife 9, e60985 (2020).
Choo, B., Zhu, M., Zhao, W. J., Jia, L. T. & Zhu, Y.-A. The largest Silurian vertebrate and its palaeoecological implications. Sci. Rep. 4, 5242 (2014).
Giles, S., Friedman, M. & Brazeau, M. D. Osteichthyan-like cranial conditions in an Early Devonian stem gnathostome. Nature 520, 82–85 (2015).
Zhu, M. et al. A Silurian placoderm with osteichthyan-like marginal jaw bones. Nature 502, 188–193 (2013).
Zhu, M. et al. A Silurian maxillate placoderm illuminates jaw evolution. Science 354, 334–336 (2016).
Gross, W. Lophosteus superbus Pander: zähne, zahnknochen und besondere schuppenformen. Lethaia 4, 131–152 (1971).
Zhu, M. et al. The oldest articulated osteichthyan reveals mosaic gnathostome characters. Nature 458, 469–474 (2009).
Choo, B. et al. A new osteichthyan from the late Silurian of Yunnan, China. PLoS ONE 12, e0170929 (2017).
Cui, X. D., Qiao, T. & Zhu, M. Scale morphology and squamation pattern of Guiyu oneiros provide new insights into early osteichthyan body plan. Sci. Rep. 9, 4411 (2019).
Chen, D. Lungfish-like antero-labial tooth addition and amphibian-like enameloid-enamel transition in the coronoid of a Devonian stem actinopterygian. J. Anat. 247, 418–441 (2025).
Zhu, Y.-A. et al. The oldest complete jawed vertebrates from the early Silurian of China. Nature 609, 954–958 (2022).
Brazeau, M. D. et al. Fossil evidence for a pharyngeal origin of the vertebrate pectoral girdle. Nature 623, 550–554 (2023).
Vaškaninová, V. et al. Marginal dentition and multiple dermal jawbones as the ancestral condition of jawed vertebrates. Science 369, 211–216 (2020).
Goujet, D. F. Les poissons placodermes du Spitsberg. Arthrodires Dolichothoraci de la Formation de Wood Bay (Dévonien inférieur) Vol. 15 (Editions Centre National Recherche Scientifique, Cahiers de Paléontologie, 1984).
Zhu, M., Yu, X. B. & Janvier, P. A primitive fossil fish sheds light on the origin of bony fishes. Nature 397, 607–610 (1999).
Gardiner, B. G. The relationships of the palaeoniscid fishes, a review based on new specimens of Mimia and Moythomasia from the Upper Devonian of Western Australia. Bull. Br. Mus. (Nat. Hist.) Geol. 37, 173–428 (1984).
Andrews, S. M., Long, J. A., Ahlberg, P. E., Barwick, R. & Campbell, K. S. W. The structure of the sarcopterygian Onychodus jandemarrai n. sp. from Gogo, Western Australia: with a functional interpretation of the skeleton. Trans. R. Soc. Edinb. Earth Sci. 96, 197–307 (2005).
Giles, S., Feilich, K., Warnock, R. C. M., Pierce, S. E. & Friedman, M. A Late Devonian actinopterygian suggests high lineage survivorship across the end-Devonian mass extinction. Nat. Ecol. Evol. 7, 10–19 (2022).
Rücklin, M. et al. Development of teeth and jaws in the earliest jawed vertebrates. Nature 491, 748–751 (2012).
Jarvik, E. Basic Structure and Evolution of Vertebrates, Vol. 1. (Academic, 1980).
Forey, P. L. History of the Coelacanth Fishes (Chapman & Hall, 1998).
Chang, M.-M. The Braincase of Youngolepis, a Lower Devonian Crossopterygian from Yunnan, Wouth-western China (University of Stockholm, Department of Geology, 1982).
Lu, J., Giles, S., Friedman, M. & Zhu, M. A new stem sarcopterygian illuminates patterns of character evolution in early bony fishes. Nat. Commun. 8, 1932 (2017).
King, B., Qiao, T., Lee, M. S. Y., Zhu, M. & Long, J. A. Bayesian morphological clock methods resurrect placoderm monophyly and reveal rapid early evolution in jawed vertebrates. Syst. Biol. 66, 599–516 (2017).
Maisey, J. G. Braincase of the Upper Devonian shark Cladodoides wildungensis (Chondrichthyes, Elasmobranchii), with observations on the braincase in early chondrichthyans. Bull. Am. Mus. Nat. Hist. 288, 1–103 (2005).
Pradel, A., Dearden, R. P., Cuckovic, A., Mansuit, R. & Janvier, P. in Ancient Fishes and their Living Relatives: a Tribute to John G. Maisey (eds Pradel, A. et al.) 183–192 (Dr. Friedrich Pfeil, 2021).
Maisey, J. G. et al. in Evolution and Development of Fishes (eds Johanson, Z. et al.) 87–109 (Cambridge Univ. Press, 2019).
Igielman, B. et al. The lower jaw of Devonian ray-finned fishes (Actinopterygii): anatomy, relationships, and functional morphology. Anat. Rec. https://doi.org/10.1002/ar.70005 (2025).
Choo, B. Revision of the actinopterygian genus Mimipiscis (=Mimia) from the Upper Devonian Gogo Formation of Western Australia and the interrelationships of the early Actinopterygii. Earth Environ. Sci. Trans. R. Soc. Edinb. 102, 77–104 (2012).
Zhu, M., Yu, X., Wang, W. & Jia, L. A primitive fish provides key characters bearing on deep osteichthyan phylogeny. Nature 441, 77–80 (2006).
Chen, D., Janvier, P., Ahlberg, P. E. & Blom, H. Scale morphology and squamation of the Late Silurian osteichthyan Andreolepis from Gotland, Sweden. Hist. Biol. 24, 411–423 (2012).
Lu, J. et al. The oldest actinopterygian highlights the crypticearly history of the hyperdiverse ray-finned fishes. Curr. Biol. 26, 1602–1608 (2016).
Schultze, H. P. & Cumbaa, S. L. in Major Events in Early Vertebrate Evolution: Palaeontology, Phylogeny, Genetics and Development 1st edn (ed. Ahlberg, P. E.) 315–332 (Taylor and Francis, 2001).
Burrow, C. J., Young, G. C. & Lu, J. Dermal skeleton of the stem osteichthyan Ligulalepis from the Lower Devonian of New South Wales (Australia). Span. J. Palaeontol. 38, 23–36 (2023).
Lu, J. Vayu 1.0, a new set of tools for visualizing surface meshes. Vert. PalAsiat. 61, 71–80 (2023).
Cui, X. D., Friedman, M., Yu, Y. L., Zhu, Y.-A. & Zhu, M. Bony-fish-like scales in a Silurian maxillate placoderm. Nat. Commun. https://doi.org/10.1038/s41467-023-43557-9 (2023).
Lu, J. et al. Megamastax supplemental data. figshare https://figshare.com/s/af44487f1dab980642a0 (2025).
