deMenocal, P. et al. Abrupt onset and termination of the African Humid Period: rapid climate responses to gradual insolation forcing. Quat. Sci. Rev. 19, 347–361 (2000).
Gasse, F. Hydrological changes in the African tropics since the Last Glacial Maximum. Quat. Sci. Rev. 19, 189–211 (2000).
Tierney, J. E., Pausata, F. S. R. & deMenocal, P. B. Rainfall regimes of the Green Sahara. Sci. Adv. 3, e1601503 (2017).
Swingedouw, D. et al. Early warning from space for a few key tipping points in physical, biological, and social-ecological systems. Surv. Geophys. 41, 1237–1284 (2020).
Crocker, A. J. et al. Astronomically controlled aridity in the Sahara since at least 11 million years ago. Nat. Geosci. 15, 671–676 (2022).
Shanahan, T. M. et al. The time-transgressive termination of the African Humid Period. Nat. Geosci. 8, 140–144 (2015).
Lézine, A.-M., Hély, C., Grenier, C., Braconnot, P. & Krinner, G. Sahara and Sahel vulnerability to climate changes, lessons from Holocene hydrological data. Quat. Sci. Rev. 30, 3001–3012 (2011).
Quade, J. et al. Megalakes in the Sahara? A review. Quat. Res. 90, 253–275 (2018).
Ghienne, J.-F., Schuster, M., Bernard, A., Duringer, P. & Brunet, M. The Holocene giant Lake Chad revealed by digital elevation models. Quat. Int. 87, 81–85 (2002).
Mohamed, A., Ahmed, E., Alshehri, F. & Abdelrady, A. The groundwater flow behavior and the recharge in the Nubian Sandstone Aquifer System during the wet and arid periods. Sustainability 14, 6823 (2022).
Kuper, R. & Kröpelin, S. Climate-controlled Holocene occupation in the Sahara: motor of Africa’s evolution. Science 313, 803–807 (2006).
Kutzbach, J. E. & Otto-Bliesner, B. The sensitivity of the African-Asian monsoonal climate to orbital parameter changes for 9000 years B.P. in a low-resolution general circulation model. J. Atmos. Sci. 39, 1177–1188 (1982).
Brovkin, V., Claussen, M., Petoukhov, V. & Ganopolski, A. On the stability of the atmosphere-vegetation system in the Sahara/Sahel region. J. Geophys. Res. Atmos. 103, 31613–31624 (1998).
Claussen, M. & Gayler, V. The greening of the Sahara during the mid-Holocene: results of an interactive atmosphere-biome model. Glob. Ecol. Biogeogr. Lett. 6, 369–377 (1997).
Braconnot, P., Joussaume, S., Marti, O. & de Noblet, N. Synergistic feedbacks from ocean and vegetation on the African monsoon response to mid-Holocene insolation. Geophys. Res. Lett. 26, 2481–2484 (1999).
Braconnot, P. et al. Results of PMIP2 coupled simulations of the Mid-Holocene and Last Glacial Maximum – part 2: feedbacks with emphasis on the location of the ITCZ and mid- and high latitudes heat budget. Clim. Past 3, 279–296 (2007).
Gasse, F. & Van Campo, E. Abrupt post-glacial climate events in West Asia and North Africa monsoon domains. Earth Planet. Sci. Lett. 126, 435–456 (1994).
Servant, M. & Servant-Vildary S. in The Sahara and the Nile (eds Williams, M. A. J. & Faure, H.) 133–162 (Balkema, 1980).
Defrance, D. et al. Consequences of rapid ice sheet melting on the Sahelian population vulnerability. Proc. Natl Acad. Sci. USA 114, 6533–6538 (2017).
Ben-Yami, M. et al. Impacts of AMOC collapse on monsoon rainfall: a multi-model comparison. Earths Future 12, e2023EF003959 (2024).
Grenier, C., Paillou, P. & Maugis, P. Assessment of Holocene surface hydrological connections for the Ounianga lake catchment zone (Chad). C. R. Geosci. 341, 770–782 (2009).
Kröpelin, S. et al. Climate-driven ecosystem succession in the Sahara: the past 6000 years. Science 320, 765–768 (2008).
Francus, P. et al. Varved sediments of Lake Yoa (Ounianga Kebir, Chad) reveal progressive drying of the Sahara during the last 6100 years. Sedimentology 60, 911–934 (2013).
Gourgaud, A. & Vincent, P. M. Petrology of two continental alkalineintraplate series at Emi Koussi volcano, Tibesti, Chad. J. Volcanol. Geotherm. Res. 129, 261–290 (2004).
Hoelzmann, P. et al. Mid-Holocene extreme precipitation in the Tibesti, Central Sahara. Nat. Commun. 16, 7426 (2025).
Sylvestre, F. et al. Decadal-scale droughts disrupted the African Humid Period in the Sahara. Zenodo https://doi.org/10.5281/zenodo.13912623 (2026).
Hoelzmann, P. et al. Mid-Holocene land-surface conditions in northern Africa and the Arabian Peninsula: a data set for the analysis of biogeophysical feedbacks in the climate system. Global Biogeochem. Cycles 12, 35–51 (1998).
Schuster, M. et al. Holocene Lake Mega-Chad palaeoshorelines from space. Quat. Sci. Rev. 24, 1821–1827 (2005).
Sylvestre, F. et al. in Le Tchad des Lacs. Les Zones Humides Sahéliennes au Défi du Changement Global (eds Raimond, C., Sylvestre, F., Zakinet, D. & Abderamane, M.) 53–64 (IRD, 2019).
Van der Meeren, T. et al. A predominantly tropical influence on late Holocene hydroclimate variation in the hyperarid central Sahara. Sci. Adv. 8, eabk1261 (2022).
Creutz, M., Van Bocxlaer, B., Abderamane, M. & Verschuren, D. Recent environmental history of the desert oasis lakes at Ounianga Serir, Chad. J. Paleolimnol. 55, 167–183 (2016).
Maley, J. & Verney, R. Populations and climatic evolution in north tropical Africa from the end of the Neolithic to the dawn of the modern era. Afr. Archaeol. Rev. 32, 179–232 (2015).
Fleitmann, D. et al. Evidence for a widespread climatic anomaly at around 9.2 ka before present. Paleoceanography 23, PA1102 (2008).
Manning, K. & Timpson, A. The demographic response to Holocene climate change in the Sahara. Quat. Sci. Rev. 101, 28–35 (2014).
Fleitmann, D. et al. Holocene ITCZ and Indian monsoon dynamics recorded in stalagmites from Oman and Yemen (Socotra). Quat. Sci. Rev. 26, 170–188 (2007).
Sereno, P. C. et al. Lakeside cemeteries in the Sahara: 5000 years of Holocene population and environmental change. PLoS One 3, e2995 (2008).
Cremaschi, M. et al. Takarkori rock shelter (SW Libya): an archive of Holocene climate and environmental changes in the central Sahara. Quat. Sci. Rev. 101, 36–60 (2014).
Kobashi, T. et al. Volcanic influence on centennial to millennial Holocene Greenland temperature change. Sci. Rep. 7, 1441 (2017).
Rohling, E. J. & Pälike, H. Centennial-scale climate cooling with a sudden cold event around 8,200 years ago. Nature 434, 975–979 (2005).
Parker, S. E. & Harrison, S. P. The timing, duration and magnitude of the 8.2 ka event in global speleothem records. Sci. Rep. 12, 10542 (2022).
Thomas, E. R. et al. The 8.2 ka event from Greenland ice cores. Quat. Sci. Rev. 26, 70–81 (2007).
Liu, Z. et al. Transient simulation of last deglaciation with a new mechanism for Bolling–Allerod warming. Science 325, 310–314 (2009).
Goosse, H. et al. Description of the Earth system model of intermediate complexity LOVECLIM version 1.2. Geosci. Model. Dev. 3, 603–633 (2010).
Ayache, M., Swingedouw, D., Mary, Y., Eynaud, F. & Colin, C. Multi-centennial variability of the AMOC over the Holocene: a new reconstruction based on multiple proxy-derived SST records. Glob. Planet. Change 170, 173–183 (2018).
Caesar, L., McCarthy, G. D. & Rahmstorf, S. Current Atlantic Meridional Overturning Circulation weakest in last millennium. Nat. Geosci. 14, 118–120 (2021).
Intergovernmental Panel on Climate Change (IPCC). Climate Change 2021: The Physical Science Basis. Contribution of Working Group I to the Sixth Assessment Report of the Intergovernmental Panel on Climate Change (eds Masson-Delmotte, V. et al.) (Cambridge Univ. Press, 2021).
Drijfhout, S., Angevaare, J. R., Mecking, J., van Westen, R. M. & Rahmstorf, S. Shutdown of northern Atlantic overturning after 2100 following deep mixing collapse in CMIP6 projections. Environ. Res. Lett. 20, 094062 (2025).
Mahamoud, A. H. Geologie und Hydrogeologie des Erdis-Beckens, NE-Tschad. In Berliner Geowissenschaftliche Abhandlungen A/76 (Dietrich Reimer Verlag, 1986).
Wolff, J. P. Carte géologique de la République du Tchad au 1/1.500.000e (BRGM Paris, 1964).
Elsheikh, A. A., Abdelsalam, M. G. & Mickus, K. Geology and geophysics of the West Nubian Paleolake and the Northern Darfur Megalake (WNPL–NDML): implication for groundwater resources in Darfur, northwestern Sudan. J. Afr. Earth Sci. 61, 82–93 (2011).
Gossel, W., Ebraheem, A. M. & Wycisk, P. A very large scale GIS-based groundwater flow model for the Nubian sandstone aquifer in Eastern Sahara (Egypt, northern Sudan and eastern Libya). Hydrogeol. J. 12, 698–713 (2004).
Poulin, C. Bilans hydrologiques et étude de la recharge de trois systèmes lac-aquifère du Bassin du Lac Tchad par une approche géochimique multi traceurs (δ18O, δ2H, 36Cl, 14C). Thèse de Doctorat, Aix-Marseille Université (2019).
Nicholson, S. E. The nature of rainfall variability over Africa on time scales of decades to millenia. Glob. Planet. Change 26, 137–158 (2000).
Ministère de l’Eau et de l’Assainissement de la République du Tchad. Synthèse hydrogéologique du nord et de l’est du Tchad (2016).
Capot-Rey, R. Borkou et Ounianga, Étude de géographie régionale. Université d’Alger, Institut de Recherches Sahariennes, Mémoire 5 (1961).
Washington, R. et al. Links between topography, wind, deflation, lakes and dust: the case of the Bodélé Depression, Chad. Geophys. Res. Lett. 33, L09401 (2006).
Lézine, A.-M., Zheng, W., Braconnot, P. & Krinner, G. Late Holocene plant and climate evolution at Lake Yoa, northern Chad: pollen data and climate simulations. Clim. Past 7, 1351–1362 (2011).
Remadji, R. et al. Modern diatom calibration data from Saharan lakes for inferring hydrochemistry. J. Paleolimnol. 69, 231–248 (2023).
van de Meeren, T. et al. in Le Tchad des Lacs. Les Zones Humides Sahéliennes au Défi du Changement Global (eds Raimond, C., Sylvestre, F., Zakinet, D. & Abderamane, M.) 127–138 (IRD, 2019).
Vandenberghe, J. Grain size of fine-grained windblown sediment: a powerful proxy for process identification. Earth Sci. Rev. 121, 18–30 (2013).
Appleby, P. G. in Tracking Environmental Change Using Lake Sediments: Basin Analysis, Coring, and Chronological Techniques (eds Last, W. M. & Smol, J. P.) 171–203 (Kluwer, 2001).
Rethemeyer, J. et al. Status report on sample preparation facilities for 14C analysis at the new CologneAMS center. Nucl. Instrum. Methods Phys. Res. B 294, 168–172 (2013).
Uchikawa, J., Popp, B. N., Schoonmaker, J. E. & Xu, L. Direct application of compound-specific radiocarbon analysis of leaf waxes to establish lacustrine sediment chronology. J. Paleolimnol. 39, 43–60 (2008).
Bronk Ramsey, C. Bayesian analysis of radiocarbon dates. Radiocarbon 51, 337–360 (2009).
Reimer, P. et al. The IntCal20 Northern Hemisphere radiocarbon age calibration curve (0–55 cal kBP). Radiocarbon 62, 725–757 (2020).
Philippsen, B. The freshwater reservoir effect in radiocarbon dating. Herit. Sci. 1, 24 (2013).
Blott, S. J. & Pye, K. GRADISTAT: a grain size distribution and statistics package for the analysis of unconsolidated sediments. Earth Surf. Process. Landf. 26, 1237–1248 (2001).
Folk, R. L. & Ward, W. C. Brazos River bar [Texas]; a study in the significance of grain size parameters. J. Sediment. Res. 27, 3–26 (1957).
Croudace, I. W., Rindby, A. & Rothwell, R. G. ITRAX: description and evaluation of a new multi-function X-ray core scanner. Geol. Soc. Lond. Spec. Publ. 267, 51–63 (2006).
Eisele, G., Haas, K. & Liner, S. in Über Probleme der holozänen Vegetationsgeschichte Osttibets. Göttinger Geographische Abhandlungen (ed. Frenzel, B.) 165–166 (Institute of Geography of the University of Göttingen, 1994).
Nakagawa, T. et al. Dense-media separation as a more efficient pollen extraction method for use with organic sediment/deposit samples: comparison with the conventional method. Boreas 27, 15–24 (1998).
Moore, P. D., Webb, J. A. & Collinson, M. E. Pollen Analysis 2nd edn (Blackwell, 1991).
Bonnefille, R. & Riollet, G. Pollens des Savanes d’Afrique Orientale (CNRS Éditions, 1980).
El Ghazali, G. A study on the pollen flora of Sudan. Rev. Palaeobot. Palynol. 76, 99–345 (1993).
El-Ghazali, G. Pollen Flora of Qatar (Scientific and Applied Research Center, Univ. Qatar, 1991).
Gosling, W. D., Miller, C. S. & Livingstone, D. A. Atlas of the tropical West African pollen flora. Rev. Palaeobot. Palynol. 199, 1–135 (2013).
Maley, J. Contributions à l’étude du bassin tchadien. Atlas de pollens du Tchad. Bull. Jard. Bot. Natl Belg. 40, 29–48 (1970).
Schüler, L. & Hemp, A. Atlas of pollen and spores and their parent taxa of Mt Kilimanjaro and tropical East Africa. Quat. Int. 425, 301–386 (2016).
Sowunmi, M. Pollen grains of Nigerian plants. Grana 13, 145–186 (1973).
Sowunmi, M. Pollen of Nigerian plants. Grana 34, 120–141 (1995).
Lézine, A.-M., Ivory, S. J., Gosling, W. D. & Scott, L. in Quaternary Vegetation Dynamics. The African Pollen Database (eds Gosling, W., Lézine, A. M. & Scott, L.) 5–13 (CRC, 2022).
Martin, A. C. & Harvey, W. J. The Global Pollen Project: a new tool for pollen identification and the dissemination of physical reference collections. Methods Ecol. Evol. 8, 892–897 (2017).
MUPA members. Montpellier University Pollen Atlas. OSU OREME Collection (2018).
APSA Members. Australasian Pollen and Spore Atlas V1.0. Australian National University. http://apsa.anu.edu.au/ (2007).
PalDat – Palynological database. https://www.paldat.org/.
PCU Pollen Database. https://science.uct.ac.za/plant-conservation/resources-databases/pcu-pollen-database.
Djamali, M. & Cilleros, K. Statistically significant minimum pollen count in Quaternary pollen analysis; the case of pollen-rich lake sediments. Rev. Palaeobot. Palynol. 275, 104156 (2020).
White, F. The Vegetation of Africa; A Descriptive Memoir to Accompany the UNESCO/AETFAT/UNSO Vegetation Map of Africa (UNESCO, 1983).
Hély, C. et al. Holocene changes in African vegetation: trade-off between climate and water availability. Clim. Past 10, 681–686 (2014).
Vincens, H. et al. African pollen database inventory of tree and shrub pollen types. Rev. Palaeobot. Palynol. 145, 135–141 (2007).
Wickens, G. E. A study of Acacia albida. Kew Bull. 23, 181–202 (1969).
Walter, H. & Breckle, S. Ökologie der Erde. Band 1. Ökologische Grundlagen in globaler Sicht. 2. Auflage (Gustav Fischer Verlag, 1991).
Battarbee, R. W., Jones, V. J., Flower, R. J., Cameron, N. G. & Bennion, H. in Tracking Environmental Change Using Lake Sediments (eds Smol J. P., Birks H. J. B. & Last, W. M.) 155–202 (Kluwer, 2001).
Krammer, K. & Lange-Bertalot, H. Süsswasserflora von Mitteleuropa. Bacillariophyceae. Teil 1: Naviculaceae (Gustav Fischer Verlag, 1986).
Krammer, K. & Lange-Bertalot, H. Süsswasserflora von Mitteleuropa. Bacillariophyceae. Teil 2: Bacillariaceae, Epithemiaceae, Surirellaceae (Gustav Fischer Verlag, 1988).
Krammer, K. & Lange-Bertalot, H. Süsswasserflora von Mitteleuropa. Bacillariophyceae. Teil 3: Centrales, Fragilariaceae, Eunotiaceae (Gustav Fischer Verlag, 1991).
Guiry M. D. & Guiry G. M. AlgaeBase. World-wide electronic publication, University of Galway. https://www.algaebase.org (2022).
Addinsoft. XLSTAT statistical and data analysis solution. https://www.xlstat.com (2022).
Gasse, F. East African Diatoms. Taxonomy, Ecological Distribution. Bibliotheca Diatomologica, Band 11 (J. Cramer, 1986).
Rirongarti, R. et al. Modern diatom calibration data from Saharan lakes for inferring hydrochemistry. J. Paleolimnol. 69, 231–248 (2023).
Wolin, J. A. & Stone, J. R. in The Diatoms: Applications for the Environmental and Earth Sciences 2nd edn (eds Smol J. P. & Stoermer E. F.) (Cambridge Univ. Press, 2010).
Ter Braak, C. J. F. & Looman, C. W. N. Weighted averaging, logistic regression and the Gaussian response model. Vegetation 65, 3–11 (1986).
Birks, H. J. B., Line, J. M., Juggins, S., Stevenson, A. C. & Ter Braak, C. J. F. Diatoms and pH reconstruction. Philos. Trans. R. Soc. Lond. B 327, 263–278 (1990).
Rirongarti R. Etude des Diatomées Actuelles des Lacs du Tchad: Taxonomie, Diversité et Calibration. Thèse de Doctorat, Université d’Aix-Marseille (2019).
Juggins, S. C2 User Guide. Software for Ecological and Palaeoecological Data Analysis and Visualisation (Univ. Newcastle, 2003).
Kaufman, D. et al. A global database of Holocene paleotemperature records. Sci. Data 7, 115 (2020).
Jomelli, V. et al. In-phase millennial-scale glacier changes in the tropics and North Atlantic regions during the Holocene. Nat. Commun. 13, 1419 (2022).
Shi, X. et al. Simulated stable water isotopes during the mid-Holocene and pre-industrial periods using AWI-ESM-2.1-wiso. Geosci. Model. Dev. 16, 5153–5178 (2023).
Danilov, S., Sidorenko, D., Wang, Q. & Jung, T. The finite-volume sea ice–ocean model (FESOM2). Geosci. Model. Dev. 10, 765–789 (2017).
Scholz, P. et al. Assessment of the Finite-volumE Sea ice-Ocean Model (FESOM2.0) – part 1: description of selected key model elements and comparison to its predecessor version. Geosci. Model. Dev. 12, 4875–4899 (2019).
Sidorenko, D. et al. Evaluation of FESOM2.0 coupled to ECHAM6.3: preindustrial and HighResMIP simulations. J. Adv. Model. Earth Syst. 11, 3794–3815 (2019).
Stevens, B. et al. Atmospheric component of the MPI-M Earth system model: ECHAM6. J. Adv. Model. Earth Syst. 5, 146–172 (2013).
Giorgetta, M. A. et al. Climate and carbon cycle changes from 1850 to 2100 in MPI-ESM simulations for the Coupled Model Intercomparison Project phase 5. J. Adv. Model. Earth Syst. 5, 572–597 (2013).
Raddatz, T. J. et al. Will the tropical land biosphere dominate the climate–carbon cycle feedback during the twenty-first century? Clim. Dyn. 29, 565–574 (2007).
Reick, C. H., Raddatz, T., Brovkin, V. & Gayler, V. Representation of natural and anthropogenic land cover change in MPI-ESM. J. Adv. Model. Earth Syst. 5, 459–482 (2013).
Vorrath, M. E. et al. Sea ice dynamics in the Bransfield Strait, Antarctic Peninsula, during the past 240 years: a multi-proxy intercomparison study. Clim. Past 16, 2459–2483 (2020).
Kageyama, M. et al. The PMIP4 Last Glacial Maximum experiments: preliminary results and comparison with the PMIP3 simulations. Clim. Past 17, 1065–1089 (2021).
Otto-Bliesner, B. L. et al. Large-scale features of last interglacial climate: results from evaluating the lig127k simulations for the Coupled Model Intercomparison Project (CMIP6)–Paleoclimate Modeling Intercomparison Project (PMIP4). Clim. Past 17, 63–94 (2021).
Hossain, A. et al. The impact of different atmospheric CO2 concentrations on large scale Miocene temperature signatures. Paleoceanogr. Paleoclimatol. 38, e2022PA004438 (2023).
Shi, X., Lohmann, G., Sidorenko, D. & Yang, H. Early-Holocene simulations using different forcings and resolutions in AWI-ESM. Holocene 30, 996–1015 (2020).
Köhler, P. et al. A 156 kyr smoothed history of the atmospheric greenhouse gases CO2, CH4, and N2O and their radiative forcing. Earth Syst. Sci. Data 9, 363–387 (2017).

